The transformers were slaves created by the Quintesson. Their bodies come from the planet dubbed Cybertron's mechanical outer shell/crust and their power cores or "sparks" are mined from the planet's volatile inner energy. "Cybertron" itself is a powerful, grouchy, sapient mechanical planet who takes exception to this mutilation and theft. Spark Eaters are the planet's defense against these parasites and a way to return its stolen life energy from the abominations burrowing into its body. The Matrix of Leadership was created from an integral part of Cybertron's core with the two fold purpose of both weakening the planet and controlling the rapidly advancing development of their slaves. The Prime is simply the lead slave who the other slaves are programmed to obey through the control of the Matrix. However the Quintesson did not count on a Prime itself using its power to drive them off and after doing so the transformers were free to grow, adapt, and advance on their own.
Eons pass and the slaves have split and war upon themselves, ultimately leaving a dormant planet behind and relatively unguarded.
Autobot and Decepticon alike are rapidly recalled to Cybertron to protect their home from the Quintessons who have ripped Wells- great holes- into the planet to mine its remaining energy. Though unresponsive and powerless itself, the planet's natural defenses rise greater then ever to fight the invaders and the abominations on its surface. Refusing to lose the planet once more the Quintessons command the newest Prime heed their demand and obey or be destroyed by the swarming Spark Eaters. It is his function, his purpose to compel the slaves to obedience, a function that he has only half fulfilled.
Horrified at the truth, that their home is a tortured and imprisoned being that their very lives were stolen from, that by existing he has taken his Autobot's true choice away, and separated, backed up to the edge of the largest Well by a tide of Spark Eaters and the Quintessons over head dangling his survival as threat and bribe, Optimus does the only thing he can.
This is the universe Orion Pax is expelled into when Cybertron spits him back out of the well. A universe where Autobot, Neutral, and Decepticon alike have no memory of Optimus Prime.
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